1992 …2020

Research output per year

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Personal profile

Research interests

My research is very applied, and is usually carried out in collaboration with farmers and/or the agricultural industries, with the aim of developing improved, welfare friendly, sustainable methods of farming. As a principle investigator I have held a large number of major grants (> £100,000) covering a wide range of projects.




These have included improving the housing systems and handling of laying hens, investigations into improving transport conditions for equidae, sheep, cattle, pigs and both laying hens and broiler (meat) chickens. The figure is from some of my work in Chile, looking at stocking densities in cattle transported by road.

I've also held major grants which have investigated the control of lameness in dairy cattle, the effects of differences in water quality on farmed trout, the control of mastitis in dairy cattle, the extent and causes of lameness in broiler chickens and the extent and causes of fin damage in farmed trout. The figure below shows one of the measurement scales we developed to assess the extent of fin erosion in farmed rainbow trout.




Other areas of investigation and interest have included surveys to assess levels of contaminants within the food chain in fish, wild collected food and in vegetables and research into campylobacter infection in broiler flocks. I’ve also had an interest in developing more humane methods of slaughter and have worked with colleagues to develop instantaneous electrical stun/kill systems for farmed fish which are now widely used throughout the UK where previously fish were left to die slowly over a matter of minutes in air or in an ice slurry. I’ve long held an interest in cetaceans and my research into the ineffectiveness of the Norwegian grenade harpoon, based on data collected from the Norwegian minke whale hunt, was adopted and presented at the annual International Whaling Commission meeting by the British government delegation. The figure below summarises the main findings of the work and shows the small target area of the whale which needs to be struck to begin to be sure of a humane kill.

Quantifying the ineffectiveness of the Norwegian harpoon grenade

The following is a list of current research projects:-

  • Humane slaughter of farmed fish
  • Fitness to travel of end-of-lay hens
  • Anaesthesia and analgesia techniques applicable to fish
  • Investigation of rearing environment and its association with feather pecking in free-range pullets
  • A review of equine welfare in England & Wales
  • The impact of keel bone fractures on the welfare of laying hens
  • Evaluation of the sustainability of UK sheep transport and marketing systems


My latest research project aims to evaluate the effects that the different types of transport and marketing systems used for sheep in the UK have on the animals. It also aims to quantify the extent to which the different systems are in operation within the UK and using a life cycle analysis approach, investigate the sustainability of the systems.

I have published over 100 articles in refereed scientific journals which cover the work described above and more. Many of the references to my work can be found using the links within this micro site, however, a search of Google Scholar using 'Knowles TG' will provide a fuller list with links to the source publications, some of which are in the public domain.



Further information about Professor Toby Knowles can be found here.

Structured keywords and research groupings

  • Cabot Institute Food Security Research

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Research Output

Analysis of risk factors for dermatitis and cellulitis in turkeys using abattoir post mortem data

Neroli, S., Prestmo, P. G., Sheard, P., Knowles, T. G. & van Klink, E. G. M., 3 Jul 2020, In : British Poultry Science. p. 1-7 7 p.

Research output: Contribution to journalArticle (Academic Journal)

  • Comparing paracetamol/codeine and meloxicam for postoperative analgesia in dogs: a non-inferiority trial

    Pacheco, M., Knowles, T. G., Hunt, J., Slingsby, L., Taylor, P. & Murrell, J., 3 Jan 2020, In : Veterinary Record. 9 p.

    Research output: Contribution to journalArticle (Academic Journal)

    Open Access
  • 78 Downloads (Pure)

    Halal food marketing: An evaluation of UK Halal standards

    Fuseini, A. & Knowles, T. G., 9 Apr 2020, (Accepted/In press) In : Journal of Islamic Marketing. 26 p.

    Research output: Contribution to journalArticle (Academic Journal)

  • Datasets

    dog osteoarthritis project

    Jenkins, H. (Creator), Hunt, J. R. (Creator), Murrell, J. (Creator), Goff, M. (Creator), Mendl, M. T. (Creator), Whay, H. (Creator) & Knowles, T. G. (Creator), University of Bristol, 21 Jan 2016



    Astra Zeneca 2015 CEO Awards - Exceptional Delivery

    Knowles, Toby G (Recipient), 25 Feb 2015

    Prize: Prizes, Medals, Awards and Grants

  • Council for Science and Animal Welfare – Global 3Rs Award

    Knowles, Toby G (Recipient), 23 Jan 2015

    Prize: Prizes, Medals, Awards and Grants

  • Fellow of the Royal Society of Biology

    Knowles, Toby G (Recipient), 1 Apr 2016

    Prize: Election to learned society

  • Activities

    • 3 - Research and Teaching at External Organisation
    • 2 Advisory work for/on national or international committee or working group
    • 2 Other

    The Research Council of Norway (External organisation)

    Toby G Knowles (Reviewer)

    19 Oct 201620 Oct 2016

    Activity: Membership typesAdvisory work for/on national or international committee or working group

    Collaborative work on animal welfare - publication

    Toby G Knowles (Contributor)

    Jul 20132014

    Activity: Other activity typesOther

    Hosting external PhD student - animal welfare

    Toby G Knowles (Contributor)

    Jul 2013Dec 2013

    Activity: Other activity typesOther

    Supervised Work

    A mouse model of haemoplasma infection, in vitro cultivation of haemoplasmas and steps towards better diagnosis of and vaccination against haemoplasmosis

    Author: Filler, S., 23 Jan 2020

    Supervisor: Helps, C. (Supervisor), Bailey, M. (Supervisor), Tasker, S. (Supervisor), Hill, D. (Supervisor) & Knowles, T. G. (Supervisor)

    Student thesis: Doctoral ThesisDoctor of Philosophy (PhD)


    Animal Welfare Aspects of Mechanical Stunning and Killing of Neonate Farm Animals and Mechanical Stunning of Adult Cattle

    Author: Grist, A., 12 May 2020

    Supervisor: Knowles, T. (Supervisor)

    Student thesis: Doctoral ThesisDoctor of Philosophy (PhD)


    The development and practical implementation of Single Pulse Ultra-High Current for humane and Halal compliant slaughter of cattle

    Author: Fuseini, A., 28 Nov 2019

    Supervisor: Knowles, T. G. (Supervisor) & Grist, A. (Supervisor)

    Student thesis: Doctoral ThesisDoctor of Philosophy (PhD)