Personal profile
Research interests
My research project entails the anatomical and histological study of dermoskeletal and endoskeletal elements among the stem-gnathostomes, a spectrum of jawless and jawed vertebrates that represent the first vertebrates to possess skeletal tissues and which record the sequential assembly of the entire vertebrate skeleton.
Methods of analysis include using synchrotron radiation X-ray tomographic microscopy (SRXTM) to obtain high-resolution data from fossil remains, applying computed tomography to interrogate complex fossil remains, employing comparative anatomy to glean evolutionary insights from the remains of fossil and living organisms, and interpreting these results using state-of-the-art model-based phylogenetic and phylogenetic-comparative methods.
I am also interested in integrating fossil data with developmental biology and gene regulation to gain a comprehensive understanding of the evolution of mineralized skeletons in vertebrates.
Supervised by Prof. Philip Donoghue and Prof. Davide Pisani.
Keywords
- vertebrate
- dermal skeleton
- endoskeleton
- evolutionary biology
- fossil fishes
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Collaborations and top research areas from the last five years
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A New Dayongaspid Galeaspid from the Silurian of the Lower Yangtze Region: Implications for Biogeography and the Evolution of Key Adaptations in Galeaspids
Zhang, Y., Shan, X., Lin, X., Gai, Z. & Donoghue, P. C. J., 21 Feb 2026, In: Journal of Earth Science. 37, 1, p. 303-316 14 p.Research output: Contribution to journal › Article (Academic Journal) › peer-review
Open AccessFile5 Downloads (Pure) -
The dermal skeleton of stem-actinopterygian Moythomasia durgaringa and its implications for the nature of the ancestral osteichthyan
Shan, X., Pape, E., Keating, J. N., Rucklin, M., Pisani, D. & Donoghue, P. C. J., 19 Mar 2026, In: Journal of Morphology. 287, 3, 24 p., e70120.Research output: Contribution to journal › Article (Academic Journal) › peer-review
Open Access -
The Galeaspida and new fossil records in China
Gai, Z. K., Zhu, M., Donoghue, P. C. J., Zhao, W., Shan, X., Lin, X. & Zhang, Y., 1 Oct 2025, Shanghai Scientific and Technical Publishers. 223 p. (Selected Studies of Palaeontology in China)Research output: Book/Report › Authored book
Projects
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