A synthetic approach to the construction of stimuli-responsive proteinosome membranes

  • Mary E Jenkinson-Finch

Student thesis: Doctoral ThesisDoctor of Philosophy (PhD)

Abstract

A major challenge facing bottom-up synthetic biology research is the construction of
protocell populations able to respond to external signals, creating functional
protocellular ecosystems with emergent behaviours. The aim of this thesis is to
construct stimuli-responsive protocell models (proteinosomes) via chemical
programming of the crosslinked membrane as a step towards this.
The polyethylene glycol (PEG) based crosslinker traditionally used in
proteinosome synthesis has the potential to be easily functionalised with the addition
of responsive moieties. Chapter 2 presents the synthesis and characterisation of two
novel stimuli-responsive crosslinkers with embedded responsive moieties and
terminating N-hydroxy succinimide (NHS) groups for protein crosslinking. One cleaves
in response to UV light, and the other in response to a decrease in pH. The synthetic
routes and full characterisation are presented and crosslinker cleavage under a variety
of conditions is described.
The choice was made to focus research into light-responsive proteinosomes
due to ease of synthesis and higher potential with regards to disassembly control.
Chapter 3 outlines the synthesis and characterisation of chemically programmed lightresponsive proteinosomes. A UV laser in a confocal microscope system is employed
causing complete disassembly of the proteinosome membranes. Tuneable
parameters, laser power and laser scan speed, are investigated as methods to control
disassembly. Confocal software-controlled positioning systems allow targeting of
individual proteinosomes, and bespoke 2D patterning within proteinosome populations
in a simpler method not possible for other phenotypes of proteinosome.
Chapter 4 presents an in-depth analysis of the light-responsive proteinosome
membrane. A novel mathematical model to describe the inherent membrane pores is
produced, agreeing with experiments determining membrane molecular weight cutoff. An investigation light-induced selective release based on molecular weight of
substrates is shown with the permeability constants of substrates estimated. These
experiments are a step towards the design and synthesis of bespoke proteinosome
membranes and developing complex functional microsystems.
Date of Award27 Sept 2022
Original languageEnglish
Awarding Institution
  • University of Bristol
SupervisorStephen Mann (Supervisor)

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